Borthwick feels Hartley's pain after Lions snub

LONDON ENGLAND- MARCH 11 Scotland players look dejected after the RBS Six Nations match between England and Scotland at Twickenham Stadium

But by opting for an expanded 41-man squad, Gatland was able to make room for England's rapid outside centre with dazzling footwork.

However, having looked in line for a shock call-up, Roberts - who has featured in the last two tours - was left out by a coach who has brought the best out of him on the worldwide stage.

We're not so sure we'd be quite so understanding we were in Scotland's position.

"If I felt I'd only scraped into the squad and they asked me to captain, I would have said I was not in the right frame of mind to do it", he said.

"He has his points of difference of footwork and speed and the lines he ran against Scotland in the Six Nations were outstanding".

Henshaw is one of 11 Irish players who have made the cut for the squad.

So Hartley joined Robshaw, Steve Borthwick and Phil de Glanville as England captains of recent vintage not selected by the Lions.

Yet there are some significant absentees, the most notable of which sees their captain Hartley overlooked in favour of his worldwide understudy Jamie George, who travels as one of three hookers alongside Rory Best and Ken Owens despite never having started a Test.

"We couldn't leave Jamie out".

"The prognosis with him is a lot more positive than was first diagnosed in terms of it's an unusual injury with a capsule thing, or something, but the prognosis and the recovery is hopefully a lot better than what was potentially first suspected".

"I'd expect we'll get a similar game-style to whatever we've got when we've played a Warren Gatland team", Hansen added.

"It wasn't the leadership and competitiveness - obviously Dylan Hartley has done a great job in terms of that for England - but if we picked Dylan and left out Jamie or Rory or Ken, we'd be asking the same questions".

"There was always going to be a tough call and someone was always going to be disappointed".

Jonny and Richie Gray were missing from Scotland's sparse two-man contingent, while Donnacha Ryan was also absent, George Kruis, Maro Itoje, Courtney Lawes, Iain Henderson and Alun Wyn Jones the preferred options.

While London Welsh were one of England's top clubs when rugby union was still an amateur sport, the professional game has not been kind to the Exiles. "Joe is unlucky and we had a discussion about the second-rows", Gatland said.

Warburton resigned as Wales captain, with lock Alun Wyn Jones leading the side in the Six Nations. He also skippered them on the successful 2013 tour of Australia.

There was criticism from Lions legend Ian McLauchlan of Gatland's decision only to select two Scots. They are the No.1 team in the world, so it's going to be a massive test for all of us going out there.

After the Lions were named, Gatland highlighted that All Black first five-eighths Beauden Barrett wasn't even kicking for the Hurricanes, emphasising that the Lions would head south with a squadron of goalkicking aces.

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